The intertidal zone (also known as the foreshore and seashore and sometimes referred to as the littoral zone) is the area that is exposed to the air at low tide and underwater at high tide (for example, the area between tide marks). This area can include many different types of habitats, including steep rocky cliffs, sandy beaches, or wetlands (e.g., vast mudflats). The area can be a narrow strip, as in Pacific islands that have only a narrow tidal range, or can include many meters of shoreline where shallow beach slope interacts with high tidal excursion.Organisms in the intertidal zone are adapted to an environment of harsh extremes. Water is available regularly with the tides but varies from fresh with rain to highly saline and dry salt with drying between tidal inundations. The action of waves can dislodge residents in the littoral zone. With the intertidal zone's high exposure to the sun the temperature range can be anything from very hot with full sun to near freezing in colder climates. Some microclimates in the littoral zone are ameliorated by local features and larger plants such as mangroves. Adaption in the littoral zone is for making use of nutrients supplied in high volume on a regular basis from the sea which is actively moved to the zone by tides. Edges of habitats, in this case land and sea, are themselves often significant ecologies, and the littoral zone is a prime example.
A typical rocky shore can be divided into a spray zone or splash zone (also known as the supratidal zone), which is above the spring high-tide line and is covered by water only during storms, and an intertidal zone, which lies between the high and low tidal extremes. Along most shores, the intertidal zone can be clearly separated into the following subzones: high tide zone, middle tide zone, and low tide zone.
Zonation
Marine biologists and others divide the intertidal region into three zones (low, middle, and high), based on the overall average exposure of the zone. The low intertidal zone, which borders on the shallow subtidal zone, is only exposed to air at the lowest of low tides and is primarily marine in character. The mid intertidal zone is regularly exposed and submerged by average tides. The high intertidal zone is only covered by the highest of the high tides, and spends much of its time as terrestrial habitat. The high intertidal zone borders on the swash zone (the region above the highest still-tide level, but which receives wave splash). On shores exposed to heavy wave action, the intertidal zone will be influenced by waves, as the spray from breaking waves will extend the intertidal region above the high tide line.
Depending on the substratum and topography of the shore, additional features may be noticed. On rocky shores, tide pools form at low tide when water is trapped in hollows. Under certain conditions, such as those at Morecambe Bay, quicksand may form.
Low tide zone (lower littoral)
This subregion is mostly submerged - it is only exposed at the point of low tide and for a longer period of time during extremely low tides. This area is teeming with life; the most notable difference with this subregion to the other three is that there is much more marine vegetation, especially seaweeds. There is also a great biodiversity. Organisms in this zone generally are not well adapted to periods of dryness and temperature extremes. Some of the organisms in this area are abalone, anemones, brown seaweed, chitons, crabs, green algae, hydroids, isopods, limpets, mussels, nudibranchs, sculpin, sea cucumber, sea lettuce, sea palms, sea stars, sea urchins, shrimp, snails, sponges, surf grass, tube worms, and whelks. Creatures in this area can grow to larger sizes because there is more available energy in the localised ecosystem and because marine vegetation can grow to much greater sizes than in the other three intertidal subregions due to the better water coverage: the water is shallow enough to allow plenty of light to reach the vegetation to allow substantial photosynthetic activity, and the salinity is at almost normal levels. This area is also protected from large predators such as large fish because of the wave action and the water still being relatively shallow.
